Tetsuro Matsuzawa is Director of the Primate Research Institute, Kyoto University, Japan. He studies chimpanzee intelligence both in the laboratory and in the wild. The Ai project began in 1978 with research on language-like skills and number concepts in a single chimpanzee, and now focuses on cultural transmission of knowledge, skills, and values across generations. Research on behavior of wild chimpanzees in their natural habitat in Guinea, West Africa since 1986 has documented use of a pair of mobile stones as hammer and anvil to crack open oil-palm nuts, and has examined hand preference, critical periods, "education by master-apprenticeship", and cultural variation across adjacent communities. Prof Matsuzawa’s many  publications include “Cognitive development in chimpanzees” (2006).